The Central U.S. Mixed Hardwoods ecoregion is among the richest in North America for herbaceous plants and shrubs, with 2,527 species (Kartesz, personal communication). The tree flora is less diverse, dominated by only a few species. The oak-hickory forest becomes more savanna-like in its northern reaches. In southern Illinois and Indiana, the forest forms a mosaic with prairie. Widepsread dominants are white oak (Quercus alba), red oak (Q. rubra), black oak (Q. velutina), bitternut hickory (Carya cordiformis), and shagbark hickory (C. ovata). Flowering dogwood (Cornus florida) often occurs in the understory, along with sassafras (Sassafras spp.) and hop hornbeam (Carpinus spp.). The shrub layer is distinct, often with evergreens, and wildflowers are common. Intact wetter sites feature American elm (Ulmus americana), tuliptree (Liriodendron tulipifera), and sweetgum (Liquidambar styraciflua).
Only about one percent of the Central U.S. Hardwoods remains as intact habitat. The majority of the ecoregion has been heavily altered by human activity, particularly conversion to agriculture, short rotation silviculture, and pasture in some areas (e.g., bluegrass).
Remaining Blocks of Habitat
All the remaining blocks of habitat in this ecoregion are smaller than 1000 km2. The most important are:
•Mammoth Cave - south-central Kentucky
•Land between the Lakes - southeastern Kentucky, northeastern Tennessee
•Hoosier National Forest, Yellowwood Nathional Forest - southern Indiana
•Edge of Appalachia - central Tennessee
•Western Highland Rim - eastern Tennessee
•Lower Missouri Ozarksm - southern Missouri
•Tennessee Cedar Glades - central Tennessee
•Missouri Ozarks/southern Mark Twain National Forest - southern Missouri, northern Arkansas
•Shawanee Hills - southern Illinois
•Big River Junction - eastern Illinois
•Wolf River - southeastern Tennessee
•Fort Campbell - northern Tennessee, southern Kentucky
Degree of Fragmentation
Most of this ecoregion has been highly fragmented, so large predators have disappeared. The remaining habitat blocks have some degree of connectivity, but few corridors exist between blocks. Degraded corridors within the blocks may be restorable, though the focus should be on improving the integrity of large blocks.
Degree of Protection
Mammoth Cave, Edge of Appalachia, and parts of the Missouri Ozarks and the Tennessee Cedar Glades are the only blocks of habitat in the ecoregion to enjoy substantial protection. These areas do not adequately protect the habitats of the central U.S. hardwoods.
Types and Severity of Threats
Urban sprawl and agricultural conversion are the greatest conversion threats to the region. Invasion of exotic grasses, cave vandalism and overuse for recreation, fire suppression in fire-maintained systems, and loss of large ungulates (bison) are degrading the remaining natural habitats. Deer poaching continues to be a problem in Kentucky and Tennessee, and collection of wild herbs is ongoing across the region.
Suite of Priority Activities to Enhance Biodiversity Conservation
•Improve integrity of existing natural habitats
•Continue to identify existing biodiversity hotspots and protect them. These are important places that conserve the full range of communities and species within the ecoregion, some of which will be small, some large, some connectible, and some not.
•Alabama Natural Heritage Program
•Arkansas Natural Heritage Commission
•Illinois Natural Heritage Division
•Indiana Natural Heritage Data Center
•Kentucky Natural Heritage Program
•Mississippi Natural Heritage Program
•Missouri Natural Heritage Database
•The Nature Conservancy
•The Nature Conservancy - Midwest Reg. Office
•The Nature Conservancy - Southeast Regional Office
•Ohio Natural Heritage Data Base
•Oklahoma Natural Heritage Inventory
•Tennessee Division of Natural Heritage
Relationship to other classification schemes
This ecoregion corresponds roughly to the southern half of Bailey’s Eastern Broadlead Forest (Continental) Province. The northern portion of Bailey’s Province should be distinguished on the basis of climate, vegetation types, and ecosystem dyanmics. Omernik classifies the Ozark Highlands as a separate ecoregion, while we believe these rightly belong with the central U.S. hardwoods.
Prepared by: A. Weakley, R. Noss, E. Dinerstein, S. Robinson, J. Adams