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Sri Lanka dry-zone dry evergreen forests (IM0212)

 

Sri Lanka dry-zone dry evergreen forests
Satellite view of the dry forest in northern Sri Lanka
Photograph by USGS


 

Where
Southern Asia: Island of Sri Lanka off the coast of India
Biome
Tropical and Subtropical Dry Broadleaf Forests

  Size
18,700 square miles (48,400 square kilometers) -- about twice the size of Vermont
Vulnerable
 
 

· Location and General Description
· Biodiversity Features
· Current Status
· Threats
· Ecoregion Justification
· References
More Photos

The Sri Lanka Dry-Zone Dry Evergreen Forests [IM0212] harbor one of Asia's largest and most viable Asian elephant (Elephas maximus) populations and a large protected area system designed specifically for elephant conservation. Therefore, the ecoregion provides one of the best opportunities to conserve Asia's largest vertebrate over the long-term. Unlike most other dry forests, the trees in the Sri Lanka Dry-Zone Dry Evergreen Forests [IM0212] retain their leaves during the dry season. Only two other ecoregions exhibit this phenology: the small East Deccan Dry Evergreen Forests [IM0204] and the Southeastern Indochina Dry Evergreen Forests [IM0210] in the Indochina bioregion. Because the evergreen dry forest ecoregion in the east Deccan Plateau is small and has lost most of its intact habitat, the Sri Lanka Dry-Zone Dry Evergreen Forests [IM0212] are the only viable example of evergreen dry forests in the bioregion.

Location and General Description

The Sri Lanka Dry-Zone Dry Evergreen Forests [IM0212] represent the tropical dry forests throughout most of the island of Sri Lanka, except for the southwestern quarter, the central mountain range, and the Jaffna Peninsula in the extreme north.

The island's geological roots date back to the Cretaceous period when, as part of the Deccan Plateau, it detached from Gondwanaland and drifted north to collide with the northern Eurasian continent about 50 million years later. Therefore, the ecoregion harbors elements of the ancient Gondwana biota. The island then became separated from the Deccan Plateau during the late Miocene about 20 million years later. Since that time, there have been several land bridges that allowed species exchanges between the island and the mainland until the final separation during the Pleistocene (Deraniyagala 1992).

The ecoregion receives about 1,500-2,000 mm of annual rainfall in the December to March northeast monsoon but is mostly dry the rest of the year. Topographically, the ecoregion is flat, except for scattered inselbergs and isolated low hills. Ritigala, a 766-m isolated peak in the central part of Sri Lanka, is the highest point between the central massif and the Western Ghats of India (Jayasuriya 1984).

Tropical dry forests cover most of the ecoregion, but within the ecoregion there are a number of distinct habitat types (of sub-regional extent). These include patches of submontane savanna and grassland-known locally as talawa-especially along the eastern and southeastern slopes of the central massif. In the northeast, lowland grasslands, locally known as villus, are associated with the floodplains of the river systems. These grasslands provide critical water and fodder for herbivores during the dry season.

Most of this ecoregion was settled and cultivated until about 500 years ago; therefore, the forest is secondary. However, several patches of old-growth forests remain and are included within protected areas (e.g., Wasgomuwa National Park and parts of Ruhuna National Park).

The evergreen dry forests are dominated by Manilkara hexandra, Chloroxylon sweitenia, Drypetes sepiaria, Feronia limonia, Vitex altissima, Syzygium spp., Drypetes sepiaria, and Chukrasia tabularis, with the scrub and regenerating forests characterized by Bauhinia racemosa, Pterospermum suberifolium, Cassia fistula, and Dichrostachys cineria (Gunatilleke and Gunatilleke 1990). Acacia thorn scrub grows in disturbed areas.

The talawa savannas are characterized by Terminalia chebula, T. belerica, Pterocarpus marsupium, Butea monosperma, Careya arborea, Anogeissus latifolia, Phyllanthus embilica, and Zizyphus spp. (McKay 1973). The dominant grasses in the villus include Cymbopogon spp., Eragrostis spp., Themeda spp., and Imperata spp. (McKay 1973).

Ritigala, the isolated hill in central Sri Lanka, is a hotspot of endemic species within this ecoregion with several endemic plants such as Madhuca clavata (Jayasuriya 1984).

Biodiversity Features

Compared with Sri Lanka's rain forests, this ecoregion does not contain very high levels of endemism. Nevertheless, it harbors one of Asia's largest elephant populations, estimated at 2,500 to 4,000 animals. The large protected area system that was designed using the elephant as a focal species provides an excellent chance for the long-term conservation of this endangered species.

The seventy-four mammal species known from the ecoregion include four near-endemic species (table 1).

Table 1. Endemic and Near-Endemic Mammal Species.

Family

Species

Cercopithecidae

Semnopithecus vetulus

Viverridae

Paradoxurus zeylonensis

Muridae

Mus fernandoni

Muridae

Vandeleuria nolthenii

An asterisk signifies that the species' range is limited to this ecoregion.

Several of the ecoregion's mammals are also listed as threatened: the endangered Asian elephant, the Sri Lankan genotype of the common leopard (Panthera pardus kotiya), and the vulnerable sloth bear (Melursus ursinus), purple-faced leaf monkey (Semnopithecus vetulus), and slender loris (Loris tardigradus) (IUCN 2000).

Bird richness is greater, with 270 species, of which nine species are near endemic (table 2).

Table 2. Endemic and Near-Endemic Bird Species.

Family

Common Name

Species

Bucconidae

Ceylon grey hornbill

Ocyceros gingalensis

Cuculidae

Red-faced malkoha

Phaenicophaeus pyrrhocephalus

Phasianidae

Ceylon junglefowl

Gallus lafayetii

Phasianidae

Ceylon spurfowl

Galloperdix bicalcarata

Turdidae

Spot-winged thrush

Zoothera spiloptera

Timaliidae

Brown-capped babbler

Pellorneum fuscocapillum

Capitonidae

Yellow-fronted barbet

Megalaima flavifrons

Psittacidae

Ceylon hanging-parrot

Loriculus beryllinus

Psittacidae

Layard's parakeet

Psittacula calthropae

An asterisk signifies that the species' range is limited to this ecoregion.

The spot-billed pelican (Pelecanus philippensis) is a globally threatened species. BirdLife International has included this entire ecoregion within an EBA, Sri Lanka (124) (Stattersfield et al. 1998).

Viable populations of the freshwater and mugger crocodiles (Crocodylus porosus and C. palustris, respectively) are found along the rivers and estuaries and have also adapted to the numerous ancient reservoirs that formed part of the extensive irrigation system. Two other large lizards, the water monitor (Varanus salvator)-the second largest lizard in the world-and the Bengal monitor (V. bengalensis), which are persecuted and hunted throughout most of their ranges, have found a safe haven in this ecoregion.

Current Status

About three-quarters of this ecoregion has been deforested; however, extensive areas of contiguous, intact forest remain in the north and north central area. Thirty-eight protected areas cover 7,842 km2, or 17 percent of the ecoregion area (table 3). This represents the largest proportion of intact forests that are included within the protected area systems of the dry forest ecoregions in the Indo-Pacific region.

Table 3. WCMC (1997) Protected Areas That Overlap with the Ecoregion.

Protected Area

Area (km2)

IUCN Category

Ruhuna (Yala) Block 1

1,030

II

Kudumbigala

40

IV

Yala East Block 1

230

II

Uda Walawe

320

II

Yala

230

I

Kataragama

10

IV

Katagamuwa

10

IV

Wirawila-Tissa

50

IV

Bundala

70

IV

Kalametiya Kalapuwa

10

IV

Maduru Oya Block 1

570

II

Nuwaragala

400

VIII

Nelugala

90

UA

Gal Oya Valley NE

120

IV

Gal Oya Valley

280

II

Sneanayake Samudra

80

IV

Gal Oya Valley SW

120

IV

Sagamam

2

IV

Lahugala Kitulana

170

II

Vavunikulam

20

IV

Madhu Road

250

IV

Kokilai Lagoon

50

IV

Giant's Tank

50

IV

Padaviya Tank

30

IV

Wilpattu Block 1

130

II

Wilpattu North

10

IV

Anaulandewa

310

VIII

Trincomalee Naval Headworks

170

IV

Great Sober Island

1

IV

Mahakandarawewa

10

IV

Anuradhapura

40

IV

Mihintale

7

IV

Hurulu

250

VIII

Seruwila-Allai

140

IV

Somawathiya Block 1

370

II

Tirikonamadu

240

IV

Ritigala

30

I

Minneriya-Giritale

60

IV

Victoria-Randenigala-Rantambe

140

IV

Floodplains

130

II

Polonnaruwa

10

IV

Minneriya-Giritale Block 1

140

IV

Wasgomuwa Lot 1

370

II

Total

6,790

 
Ecoregion numbers of protected areas that overlap with additional ecoregions are listed in brackets.

Two of these protected areas complexes-Yala/Ruhuna and Wilpattu national parks-exceed 1,000 km2 in area, and several other reserves form contiguous complexes that exceed 500 km2. Many of these larger reserves and reserve complexes contain elephant populations that can contribute significantly to a regional conservation program.

Types and Severity of Threats

The primary overarching threats are from deforestation caused by agriculture, resettlements, and small-scale logging. Encroachment into protected areas, several of which lack adequate protection and management, also poses important threats that warrant attention. Overall, however, the protected area system is extensive enough to conserve the ecoregion's biodiversity, given effective management. The cultural and religious taboos against killing and hunting wildlife-most of the country is Buddhist-have served well to protect wildlife relative to the rest of Asia.

Justification of Ecoregion Delineation

The dry zone has been commonly used as a distinct bioclimatic zone in Sri Lanka. We used MacKinnon's (1997) map of original forest cover to separate the dry evergreen forests of Sri Lanka from the wet-zone moist forests and represented the former in the Sri Lanka Dry-Zone Dry Evergreen Forests [IM0212]. This ecoregion corresponds to MacKinnon's (1997) biounit S13 and overlaps with floristic zones 1 and 2 identified by Ashton and Gunatilleke (1987).

References

References for this ecoregion are currently consolidated in one document for the entire Indo-Pacific realm.
Indo-Pacific Reference List

Prepared by: Eric D. Wikramanayake and Savithri Gunatilleke
Reviewed by:

This text was originally published in the book Terrestrial ecoregions of the Indo-Pacific: a conservation assessment from Island Press. This assessment offers an in-depth analysis of the biodiversity and conservation status of the Indo-Pacific's ecoregions.

For more general information on this ecoregion, go to the WildWorld version of this description.

All text by World Wildlife Fund © 2001